Bifurcation Analysis of a Holling–Tanner Model with Generalist Predator and Constant-Yield Harvesting

In this paper, we introduce constant-yield prey harvesting into the Holling–Tanner model with generalist predator. We prove that the unique positive equilibrium is a cusp of codimension 4. As the parameter values change, the system exhibits degenerate Bogdanov–Takens bifurcation of codimension 4. Using the resultant elimination method, we show that the positive equilibrium is a weak focus of order 2, and the system undergoes degenerate Hopf bifurcation of codimension 2 and has two limit cycles. By numerical simulations, we demonstrate that the system exhibits homoclinic bifurcation and saddle–node bifurcation of limit cycles as the parameters are varied. The main results show that constant-yield prey harvesting and generalist predator can lead to complex dynamic behavior of the model.