A multi-ootaxic assemblage from the Lower Cretaceous of the Cameros Basin (La Rioja; Northern Spain)

Here we describe the new fossil site of El Horcajo in the vicinity of Trevijano (La Rioja, Spain). It is located in the palustrine facies of the Enciso Group (Cameros Basin). This new locality has provided dozens of eggshell fragments, together with other vertebrate remains and charophyte fructifications, which allow dating of the locality as Valanginian-Hauterivian. Five ootaxa have being recognized: 1) the Spheroolithidae Guegoolithus turolensis; 2) a surprisingly thick Prismatoolithidae indet., with certain affinities to the oogenus Sankofa, that may represent a new oogenus and oospecies for this oofamily; 3) the Krokolithidae Krokolithes sp.; 4) recrystallized ?Testudoolithidae eggshells; and 5) ?Geckoolithidae eggshells of uncertain affinity. This oodiversity is similar to that of other eggshell microsites. The ootaxonomic list differs from the other Iberian locality of the same age, Pochancalo 1 (Villanueva de Huerva Formation, Zaragoza) in lacking sauropod and megalosauroid theropod eggshells, but shares the presence the ornithopod and coelurosaurian theropod eggshells with other younger microsites found in similar facies. The age of the new locality has immediate consequences for the tectono-sedimentary framework of the Cameros Basin, and the new data will need to be considered in future stratigraphical works. Furthermore, the reliability of eggshells as biostratigraphic markers is questioned, and the viability of Guegoolithus as a guide fossil for the lower Barremian is rejected