Sliding Dynamics and Bifurcations in the Extended Nonsmooth Filippov Ecosystem

We propose a nonsmooth Filippov refuge ecosystem with a piecewise saturating response function and analyze its dynamics. We first investigate some key elements to our model which include the sliding segment, the sliding mode dynamics and the existence of equilibria which are classified into regular/virtual equilibrium, pseudo-equilibrium, boundary equilibrium and tangent point. In particular, we consider how the existence of the regular equilibrium and the pseudo-equilibrium are related. Then we study the stability of the standard periodic solution (limit cycle), the sliding periodic solutions (grazing or touching cycle) and the dynamics of the pseudo equilibrium, using quantitative analysis techniques related to nonsmooth Filippov systems. Furthermore, as the threshold value is varied, the model exhibits several complex bifurcations which are classified into equilibria, sliding mode, local sliding (boundary node and focus) and global bifurcations (grazing or touching). In conclusion, we discuss the importance of the refuge strategy in a biological setting.