Most common approaches to predicting or documenting seedling emergence are imprecise. Mechanistic models that simulate seed dormancy and germination and seedling elongation as functions of measured or estimated environmental variables seem to be the most promising approach to the problem, but they also are the most difficult models to develop. These models will need to integrate soil water potential and soil temperature (hydrothermal time), diurnal soil temperature fluctuations, oxygen deficiency, light quality, and seed burial depth to better describe the direct and interactive effects on and among seed dormancy alleviation and induction, seed germination, and seedling elongation. In the meantime, creation and use of simpler empirical models, which also employ microclimate and soil factors for predictions, may provide sufficiently accurate predictions of seedling emergence until better mechanistic models are developed.